Algae

Algae
Organisms that perform oxygenic photosynthesis, except land plants
Marine algae growing on the sea bed in shallow waters
Freshwater microscopic unicellular and colonial algae
Algal divisions and phyla[1][2]
Prokaryotic	Cyanobacteriota
Eukaryotic (primary endosymbiosis)	Glaucophyta, Rhodophyta, Prasinodermophyta, Chlorophyta, Charophyta*
Eukaryotic (secondary endosymbiosis)	Chlorarachniophyta, Chromerida, Cryptophyta, Dinoflagellata, Euglenophyta, Haptophyta, Ochrophyta
*paraphyletic, it excludes land plants
Diversity
Living	50,605 species
Fossil	10,556 species

Algae (UK: /ˈælɡiː/ AL-ghee, US: /ˈældʒiː/ ^ⓘ AL-jee;^[3] sg.: alga /ˈælɡə/ ^ⓘ AL-gə) is an informal term for any organisms of a large and diverse group of photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular microalgae, such as cyanobacteria,^[a] Chlorella, and diatoms, to multicellular macroalgae such as kelp or brown algae which may grow up to 50 metres (160 ft) in length. Most algae are aquatic organisms and lack many of the distinct cell and tissue types, such as stomata, xylem, and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds. In contrast, the most complex freshwater forms are the Charophyta, a division of green algae which includes, for example, Spirogyra and stoneworts. Algae that are carried passively by water are plankton, specifically phytoplankton.

Algae constitute a polyphyletic group^[4] because they do not include a common ancestor, and although eukaryotic algae with chlorophyll-bearing plastids seem to have a single origin (from symbiogenesis with cyanobacteria),^[5] they were acquired in different ways. Green algae are a prominent example of algae that have primary chloroplasts derived from endosymbiont cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from endosymbiotic red algae, which they acquired via phagocytosis.^[6] Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction via spores.^[7]

Algae lack the various structures that characterize plants (which evolved from freshwater green algae), such as the phyllids (leaf-like structures) and rhizoids of bryophytes (non-vascular plants), and the roots, leaves and other xylemic/phloemic organs found in tracheophytes (vascular plants). Most algae are autotrophic, although some are mixotrophic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy, myzotrophy or phagotrophy. Some unicellular species of green algae, many golden algae, euglenids, dinoflagellates, and other algae have become heterotrophs (also called colorless or apochlorotic algae), sometimes parasitic, relying entirely on external energy sources and have limited or no photosynthetic apparatus.^[8][9][10] Some other heterotrophic organisms, such as the apicomplexans, are also derived from cells whose ancestors possessed chlorophyllic plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria that produce oxygen as a byproduct of splitting water molecules, unlike other organisms that conduct anoxygenic photosynthesis such as purple and green sulfur bacteria. Fossilized filamentous algae from the Vindhya basin have been dated to 1.6 to 1.7 billion years ago.^[11]

Because of the wide range of types of algae, there is a correspondingly wide range of industrial and traditional applications in human society. Traditional seaweed farming practices have existed for thousands of years and have strong traditions in East Asian food cultures. More modern algaculture applications extend the food traditions for other applications, including cattle feed, using algae for bioremediation or pollution control, transforming sunlight into algae fuels or other chemicals used in industrial processes, and in medical and scientific applications. A 2020 review found that these applications of algae could play an important role in carbon sequestration to mitigate climate change while providing lucrative value-added products for global economies.^[12]

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